Phylogenies are based on Common Ancestries 1. Distinguish between phylogeny and systematics. -evolutionary history -biological diversity Phylogenies are the evolutionary history of a species or group of related species. Systematics is the study of biological diversity in an environmental context. 2. Describe the process of sedimentation and the formation of fossils. Explain which portions of organisms are most likely to fossilize. -insulate remains -exoskeleton -sediments bury exoskeleton When the plant or animal dies and falls into the water, the water insulates the remains. Mostly the exoskeleton becomes fossilized because bacteria in the water would consume the softer body parts. During sedimentation, sediments bury the exoskeleton over time and fossilizes the animal or plant remains. 3. Explain why it is crucial to distinguish between homology and analogy before selecting characters to use in the reconstruction of phylogeny. -similarity in characteristics -shared ancestry -similarities -convergent evolution Homology is the similarity in characteristics from a shared ancestry. Analogies are the similarities between two species that is tied to convergent evolution. 4. Explain why bird and bat wings are homologous as vertebrate forelimbs but analogous as wings. -same origin -not same structurally -same function Homologous means that it evolved from the same origin; however the forelimbs of bats and birds are not the same structurally. 5. Define molecular systematics. Explain some of the problems that systematics may face in carrying out molecular comparisons of nucleic acids. -evolutionary relationships -sequences -bases -lengths Molecular systematics uses molecular data to determine evolutionary relationships. Some problems arise when some distinctly related species have many different bases and lengths. Phylogenetic Systematics: Connecting Classification with Evolutionary History 6. Explain the following characteristics of the Linnaean system of classification: a. binomial nomenclature -genus In binomial nomenclature, the genus is always capitalized and the specific epithet are never capitilizaed as well. b. hierarchical classification -comprehensive groups Species are placed into more comprehensive groups. 7. List the major taxonomic categories from most to least inclusive. -closely related -family -orders -classes -phyla -kingdoms -domains Species are grouped to the same genus based on how closely related they appear to be. It goes from families to orders to classes to phyla to kingdoms and to into domains. 8. Define a clade. Distinguish between a monophyletic clade and paraphyletic and polyphyletic groupings of species. -ancestral species -decedents -common ancestors -different species A clade is a grouping of species that include the ancestral species and its descendants. A monophyletic clade includes all common ancestors and its decedents. A paraphyletic grouping includes groups deriving from two or more species. 9. Distinguish between shared primitive characters and shared derived characters. -shared by clade -evolutional -particular clade Primitive characters are a character shared by members of a particular clade. A shared derived character is an evolutional novelty that is unique to a particular clade. 10.Explain how shared derived characters can be used to construct a phylogenetic diagram. -arose in history All of the features of an organism arose at some point in life, so it is possible to determine which clade in which each derived character appeared. 11.Explain how outgroup comparison can be used to distinguish between shared primitive characters and shared derived characters. -related to monophyletic group An outgroup comparison shows that a species or group is related to a monophyletic group. 12.Define an ingroup. -evolutionary relationship An ingroup is a species or group of species whose evolutionary relationship is what we want to determine. 13.Distinguish between a phylogram and an ultrameric tree. -character changes -root-to-tip -lineages A phylogram tree branches the character changes,while a ultrametric tree has root-to-tip path lengths for all lineages. 14.Discuss how systematists use the principles of maximum parsimony and maximum likelihood in reconstructing phylogenies. -simplest explanation -consistent facts -DNA change over time According to the principles of maximum parsimony, we should investigate the simplest explanations that are consistent with the facts. The principle of maximum likelihood states certain rules about how DNA changes over time. 15.Explain why any phylogenetic diagram represents a hypothesis about evolutionary relationships among organisms. -no way to measure Any phylogenetic diagram is viewed as a diagram because there are no ways to measure of the hypothesis is accurate or not. 16.Distinguish between orthologous and paralogous genes. Explain how gene duplication has led to families of paralogous genes. -homologous genes -gene duplication Orthologous genes are homologous genes that are found in different species because of speciation. Paralogous genes result from gene duplication. There are more than one copy found in the same genome. 17.Explain how molecular clocks are used to determine the approximate time of key evolutionary events. Explain how molecular clocks are calibrated in actual time. -absolute time of evolutionary change -graphing genetic differences A molecular clock measures the absolute time of evolutionary change, based on observations. Molecular sticks or a gene can be calibrated by graphing the numbers of genetic differences. 18.Describe some of the limitations of molecular clocks. -DNA changes -past fossil evidence Some of the limitations include DNA changes and the when the molecular clock extends over the fossil evidence. 19.Explain the neutral theory of evolutionary change. -fitness -natural selection The neutral theory of evolutionary change is that evolutionary change in genes and proteins has no effect on fitness and therefore is not effected by natural selection. 20.Explain how scientists determined the approximate time when HIV-1 M first infected humans. -comparing sequences Scientists determined the approximated time when HIV-1 M first infected humans by comparing sequences of HIV viruses from samples taken during various times during the epidemic. 21.Describe the evidence that suggests there is a universal tree of life. -ribosomal RNA -Three major categories The tree of life is based on ribosomal RNA sequences. All life on earth can be placed in one of the three major categories, which leads scientists to believe that all life started from a common ancestor.
Concept Map Words:
analogy binomial clade cladistics cladogram class domain family fossil record genus homoplasy ingroup kingdom maximum likelihood maximum parsimony molecular clock molecular systematics monophyletic neutral theory order orthologous genes outgroup paralogous genes paraphyletic phylogenetic tree phylogeny phylogram phylum polyphyletic shared derived character shared primitive character specific epithet systematics taxon taxonomy ultrametric tree
Memorize the Word Roots analog- 5 proportion (analogy: similarity due to convergence) bi- 5 two; -nom 5 name (binomial: a two-part latinized name of a species) clado- 5 branch (cladogram: a dichotomous phylogenetic tree that branches repeatedly) homo- 5 like, resembling (homology: similarity in characteristics resulting from a shared ancestry) mono- 5 one (monophyletic: pertaining to a taxon derived from a single ancestral species that gave rise to no species in any other taxa) parsi- 5 few (principle of parsimony: the premise that a theory about nature should be the simplest explanation that is consistent with the facts) phylo- 5 tribe; -geny 5 origin (phylogeny: the evolutionary history of a taxon
Chapter 26 Concept Map: 10 Points
The Tree of Life: An I ntroduction to Biological Diversity
Objectives 22. The Origin of Life 1. Describe the four stages of the hypothesis for the origin of life on Earth by chemical evolution. -nonliving synthesis -protobionts -origins -macromolecules The four stages include the abiotic synthesis of organic molecules, the joining of those organic molecules into macromolecules, packaging them into protobionts, and the origin of self-replicating molecules that make inheritance possible. 2. Describe the contributions that A. I. Oparin, J.B.S. Haldane, and Stanley Miller made toward developing a model for the abiotic synthesis of organic molecules. Describe the conditions and locations where most of these chemical reactions probably occurred on Earth. -organic molecules -conditions Oparin and Haldane discovered conditions on early earth led to the formation of small organic materials. Miller simulated conditions on early earth. 3. Describe the evidence that suggests that RNA was the first genetic material. Explain the significance of the discovery of ribozymes. -information transfer -copied abiotically -variety of catalytic functions -pre-date enzymes RNA is central to information transfer and can be copied abiotically. Ribozymes have a variety of catalytic functions, they are pre-date enzymes. 4. Describe how natural selection may have worked in an early RNA world. -stability & replicability -complementary catalytic functions -covalent linkage -amino acids RNAs with stability and replicability are favored, complementary catalytic functions allow for groups of RNA, and covalent linkages with catalytic function. 5. Describe how natural selection may have favored the proliferation of stable protobionts with self-replicating, catalytic RNA. -self-replicating -catalytic -inherited characteristics A protobiont with self-replicating, catalytic RNA would be different. If a protobiont could grow and pass on its RNA to its daughters its limited inherited characteristics could be affected by natural selection. 23. Introduction to the History of Life 6. Explain how the histories of Earth and life are inseparable. -geological events -chemical changes The history of Earth and life are inseparable because geological events affect biological evolution. Likewise organisms cause major chemical changes on Earth. 7. Explain how index fossils can be used to determine the relative age of fossil-bearing rock strata. Explain how radiometric dating can be used to determine the absolute age of rock strata. Explain how magnetism can be used to date rock strata. 8. Describe the major events in Earths history from its origin until 2 billion years ago. In particular, note when Earth first formed, when life first evolved, and what forms of life existed in each eon. -4.6 bya -3.5 bya prokaryote -2.1 bya eukaryots The Earth was formed about 4.6 billion years ago, 3.5 billion years ago prokaryotes appeared, and 2.1 billion years ago, eukaryotes evolved. 9. Describe the mass extinctions of the Permian and Cretaceous periods. Discuss a hypothesis that accounts for each of these mass extinctions. -marine animals -major volcanoes -cretaceous-comet The extinction of the Permian period killed nearly all marine animals and that was due to major volcanoes. The cretaceous period reached it end because of a comet. 24. The Major Lineages of Life 10. Describe how chemiosmotic ATP production may have arisen. 11. Describe the timing and significance of the evolution of oxygenic photosynthesis. 12. Explain the endosymbiotic theory for the evolution of the eukaryotic cell. Describe the evidence that supports this theory. -endosymbiont -inner membranes -organelles replicate -single circular DNA Some of the evidence that supports the endosymbiosis theory includes when a cell lives in a another cell, when organelles share traits with prokaryotes, when inner membranes have homologous enzymes. 13. Explain how genetic annealing may have led to modern eukaryotic genomes. 14. Describe the timing of key events in the evolution of the first eukaryotes and later multicellular eukaryotes. -4.6 bya -3.5 bya prokaryote -2.1 eukaryotes The Earth was formed about 4.6 billion years ago, 3.5 billion years ago prokaryotes appeared, and 2.1 billion years ago, eukaryotes evolved. 15. Explain how the snowball-Earth hypothesis explains why multicellular eukaryotes were so limited in size, diversity, and distribution until the late Proterozoic. -earth was frozen over The snowball-Earth hypothesis refers to the idea that some point in Earths history, the entire earth, including oceans were frozen over. 16. Describe the key evolutionary adaptations that arose as life colonized land. -reproduce on land -avoid dehydration Some key evolutionary adaptations that took place were the ability to reproduce on land and to avoid dehydration. 17. Explain how continental drift explains Australias unique flora and fauna. 18. Explain why R. H. Whittakers five-kingdom system has been replaced by a new system with three domains. -Nutritional characteristics -evolutionary similarities Whitakers system has been replaced because it grouped by nutritional characteristics and the three domains system groups by evolutionary similarties.
Concept Map Words:
colony genetic annealing geologic record half-life magnetic reversal Pangaea protobiont radiometric dating ribozyme serial endosymbiosis snowball Earth hypothesis stromatolite three-domain system
Memorize the Word Roots proto- 5 first (protobionts: aggregates of abiotically produced molecules) stromato- 5 something spread out; -lite 5 a stone (stromatolite: rocks made of banded domes of sediment in which are found the most ancient forms of life